Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

She was believed to be around 141 years old when she was euthanized due to "ongoing bone conditions related to advanced age," the zoo said Known as "the Queen of the Zoo," Gramma was in the first ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor Gramma, the Galápagos tortoise, the oldest animal at ...